Luther Burbank: The Plant Inventor

Luther Burbank was born in Massachusetts in 1849, the 13th of 15 children.  As a child, he had an interest in nature.  With an inheritance from his father, who died when Burbank was 18 years old, he purchased a 17-acre farm in Massachusetts. 

Luther Burbank, ca. 1915

Burbank bought his small farm intending to be a market farmer, selling seasonal fruits and vegetables.  He was handicapped by challenging climate and growing conditions and competition from other market farmers with a head start.  He could see that he would have to produce the best produce and offer it before his competitors could, which pushed him down the path of improving the plants he grew.

Burbank’s methods for improving plants

Burbank found his inspiration as a breeder of improved plants in his local library where he found the writings of Charles Darwin.  He could see that the concept of natural selection described by Darwin, applied equally to breeding plants.  He wrote, “It opened a new world to me.  It told me, in plain simple sentences, as matter-of-fact as though its marvelous and startling truths were commonplace that variations seemed to be susceptible, through selection, of permanent fixture in the individual…I doubt if it is possible to make anyone realize what this book meant me.” (1)

Burbank quickly put his new understanding of selection to use as a means to improve the food crops he grew.  He developed the Burbank potato that is known today as the Burbank russet potato.  He found a seed ball on his potato plants containing 23 seeds.  He grew the seeds, selecting the plants that produced the best potatoes in successive crops until he had a potato with smooth skin and few eyes that tasted good and stored well. It was also mildly resistant to the blight that caused the potato famine in Ireland, which killed one million people and caused a mass exodus from Ireland.

He understood that he had created a valuable commodity, but he couldn’t see how he could profit from it because he couldn’t produce it at scale on his small property and he didn’t have the commercial infrastructure to market it widely.  At the time, it wasn’t possible to patent new plant varieties, so he sold his new variety to an established seed merchant for $150. It was a paltry sum, even at the time, but it was the beginning of a business model that financed most of Burbank’s career as a plant inventor. 

Charles Darwin also introduced Burbank to another method of improving plants in his publication, “The Effects of Cross and Self-Fertilization in the Vegetable Kingdom.”  Burbank described how Darwin led him to the realization that hybridization is another means of improving the quality and performance of plants:  “One sentence in the very introductory chapter of that volume opened the door of my mind and took possession of my fancy.  After discussing briefly the marvel of cross- and self-fertilization in plants, Darwin said: ‘As plants are adapted by such diversified and effective means for cross-fertilization, it might have inferred from this fact alone that they derive some great advantage from the process; and it is the object of the present work to show the nature and importance of the benefits thus derived [from hybridization].’” (1)

Darwin identified natural selection and hybridization as tools of evolution that produced plants and animals best adapted to current environmental conditions.  When environmental conditions changed, as they have constantly over 4.2 billion years of the Earth’s existence, natural selection and hybridization enabled the survival of plants and animals best adapted to changed conditions. Using the same methods, but different criteria, Burbank bred the plants that conformed to the needs of humans:  the most flavorful fruit, sturdy enough to be transported from fields to tables and the most beautiful flowers, in the opinion of humans.  Burbank directed and accelerated evolution to serve humans, using the tools of natural evolution. 

Grafting the branches of one type of fruit onto the root stock of another species of tree was the third method Burbank used to create new plants.  Most orchard fruit is grown from grafting because growing fruit trees from seeds is unpredictable.  The seeds of a flavorful apple, don’t necessarily grow into a tree that produces an equally flavorful apple.  It takes years for some fruit trees to produce fruit, which can ultimately produce disappointing fruit.  Grafting is a means of reducing risk and accelerating production.

California’s Bounty

After years of economic hardship, Burbank moved to California in 1875 at the age of 26 to join his brothers.  Instantly, he was an enthusiastic promoter of the ideal climate and growing conditions of his new home in Santa Rosa. He bought 4 acres of land where he built a greenhouse, nursery, and experimental fields.  Later, he bought an 18-acre plot of land in nearby Sebastopol he called the Gold Ridge Farm, where his experiments expanded.

In California, Burbank had the climate and the acreage needed to run many experiments on fruit and nut trees as well as vegetables and flowers simultaneously.  Each experiment required planting thousands of individual plants, grown through many generations.  These experiments produced hundreds of varieties of many species of plants:

Fruits
113 plums and prunes
69 nuts
35 fruiting cactus
16 blackberries
13 raspberries
11 quinces
11 plumcots
10 cherries
10 strawberries
10 apples
8 peaches
6 chestnuts
5 nectarines
4 grapes
4 pears
3 walnuts
2 figs
1 almond
Grains, grasses, forage
9 types
Vegetables
26 types
Ornamentals
91 types
Source:  Wikipedia

Cross between Burbank and Satsuma plums

Finding His Tribe:  Scientist or Businessman?

By the turn of the century, Burbank had made his reputation as the creator of new plant varieties.  He captured the attention of scientists who wanted to adopt him into their community, learn his methods, and teach them to their students:  “The San Francisco Chronicle advocates the seizing of Luther Burbank at his home in Santa Rosa and placing him in a chair at Stanford University….The main thing is to get the recluse away from his practical experiments long enough to tell people what he has done.”  Los Angeles Times, June 7, 1901.

Burbank was invited to give a series of lectures at Stanford University at a time when botanical scientists were newly influenced by the discovery of the role of genetics in producing individual variations in plants and animals. Gregor Mendel’s studies of genetic variation done in the 1860s was buried in the archives of Mendel’s local botanical society until 1900, when they were rediscovered.

Burbank’s audience at Stanford was expecting his lecture to reflect the mechanistic determinism of Mendelian genetics.  Instead, they got a dose of Burbank’s almost mystical view of the workings of nature “…as an intricate web of vibrations and magnetic forces where ‘all motion, all life, all force, all so-called matter are following the same law of heredity found in plants and animals, a forward movement toward attraction through lines of least resistance.’” (1)

Today, our understanding of genetics is more nuanced than it was over one hundred years ago and it is more consistent with Burbank’s observations.  With the help of molecular analysis, we now know that there are hundreds of unexpressed genes that are latent unless triggered in response to specific growing conditions as well as random mutations.  Burbank’s view of variation in nature was based on close, persistent observation and his own subjective intuition, based on decades of experience.

The Carnegie Institute of Technology tried to bridge this gap between science and Burbank’s art of creating new plant varieties by giving him a generous grant of $10,000 per year on the condition that a botanical scientist would trail Burbank in the field and turn his art into a data-driven algorithm capable of replicating Burbank’s accomplishments. 

The scientist assigned to that task was immediately frustrated by the haphazard jumble of Burbank’s sketchy record-keeping.  Watching Burbank in the field was equally frustrating.  Burbank couldn’t translate the choices he made into words because his judgment was intuitive.  Finally, The Carnegie Institute lost patience with the project and terminated the grant.

Henry Ford and Thomas Edison came to visit Luther Burbank in 1915 after their visit to the Panama-Pacific Exposition in San Francisco.  It was a meeting of the minds and kindred spirits. They were businessmen whose commercial success was based on tireless effort, continuous incremental improvement, and practical invention. They were Burbank’s tribe, who became fast friends for the rest of their lives. 

Thomas Edison, Luther Burbank, Henry Ford.  Santa Rosa, 1915

Burbank Defends Evolution

It took Charles Darwin nearly 20 years to publish his treatise on evolution, On the Origin of Species, partly because he knew it challenged some of the basic premises of organized religion to which his family was committed.  The evidence that life on Earth evolved over millions of years directly contradicted the religious belief that God created all life on Earth, as it presently exists, only 6,000 years ago.  Evolution is also inconsistent with the religious belief that humans are chosen by God to rule the world and that all other creatures are subservient to our command. 

In fact, pushback to the concept of evolution was minimal in the 19th century after Origin of Species was published in 1859.  Full-throated opposition to evolution emerged in the 20th century and is epitomized by the Scopes trial that occurred in 1925, just one year before Burbank’s death. 

The Scopes trial occurred because the state of Tennessee banned the teaching of evolution in public schools. The ACLU persuaded a high school teacher, John Scopes, to test the law.  Two of the greatest orators of the time, stepped forward to try this important case.  Clarence Darrow, defended Scopes for the ACLU.  William Jennings Bryon was the prosecutor for the state of Tennessee. 

Both Darrow and Bryon asked Luther Burbank to appear as a witness at the trial, which was an indication that the public was confused about Burbank’s close relationship with the natural world.  Much to the disappointment of William Jennings Bryon, who considered himself a personal friend of Burbank’s, Burbank came down unequivocally in support of the teaching evolution.

In a letter submitted as evidence in the trial, Burbank said, “Those who would legislate against the teaching of evolution should also legislate against the teaching of gravity, electricity, and the unreasonable velocity of light, and also introduce a clause to prevent the use of the telescope, the microscope…or any other instrument of precision which in the future may be invented…for the discovery of truth.” (1)

Despite Burbank’s effort, Scopes was found guilty and the ban on teaching evolution in Tennessee remained in effect until 1967.  In 2024, the Gallup Poll reported that only 24% of Americans believe in evolution unguided by God, a percentage that has increased steadily since 2000.  The dominate view—at 37%–is that humans were created by God in their present form:

In a series of interviews with the news media, Burbank expressed his doubts about the afterlife and his admiration and kinship with Christ as a man rather than a deity:  “[Christ] was an infidel of his day because he railed against the prevailing religions and his government.  I am a lover of Christ as a man, and his work and all things that help humanity, but nevertheless just as he was an infidel then, I am an infidel today.” (1)

Burbank’s Last Success

By any measure, Burbank must be considered a success.  Although he managed to make a living, he was not wealthy because his plant inventions could not be patented.  Without patent protection, profits were realized by seed merchants, nurseries, and agricultural operations. 

Burbank made many appeals to the US Patent Office for patent protection.  His appeals sounded desperate and angry about the unfairness that often threatened him with economic ruin.  His death in 1926 at the age of 77 sparked another campaign by other plant breeders to extend patent law to the development of new plant varieties. 

The effort to extend patent law to plants was boosted by the Great Depression, which began in 1929.  Farmers are always in debt as they must borrow money to plant their next crop.  When commodity prices collapsed during the depression, many farmers lost their farms. 

The proposal to extend patent law to plant “inventions” was perceived by many politicians as a way to help farmers, although the logic of that connection is questionable because patented seeds are likely to be more expensive.  Despite that concern, the Plant Patent Act was passed with little opposition in 1930.  There were many limitations on the first patent law, many of which have since been revised.

Sixteen of Burbank’s creations received patents, a small fraction of the plants being developed at the time of Burbank’s death.

We can still learn from Luther Burbank

I encourage readers to visit Luther Burbank’s home and garden in Santa Rosa, which is now a free public park, and the museum that is open during summer months.  You will find many informational signs throughout the garden about Burbank’s inventions.  You won’t find any hint of a nativist bias in the signs.  This sign about creating a garden for butterflies makes it clear that these lovely creatures have no preference for native plants:

Luther Burbank Garden, Santa Rosa, CA. 2025

(1) The Garden of Invention:  Luther Burbank and the Business of Breeding Plants, Jane S. Smith, Penguin Books, 2009

The Light Eaters: Plants will find a way to survive…if we let them

“Life finds a way, if given a chance.” – The Light Eaters

The Light Eaters was written by Zoë Schlanger, a science journalist who covered climate change before writing Light Eaters. (1)  She explains her pivot to botanical science as a retreat from the oppressive gloom of climate change.  It proved a wise choice, as she found much to cheer us in the remarkable capabilities of plants to adapt to challenges, defend themselves against their predators and competitors, and collaborate with their plant and animal neighbors. 

Ms. Schlanger believes that botanical research has lagged behind other biological inquiry partly because of a detour unwisely taken by journalists in the 1960s and 70s that projected human traits onto plants, such as intelligence and consciousness.  Humanizing animals and plants is considered a dangerous source of bias by scientists.  When scientists described plant behavior in human terms, they were often ridiculed by their colleagues and their research projects weren’t funded.  Researchers of the capabilities of plants have been trained to avoid anthropomorphic terms to describe plant behavior.  Although Ms. Schlanger tried to observe that rule, I will give myself more leeway because most of my readers are not scientists.

Plants don’t have the mobility that enables them to fly or run away from threats.  We might think of them as handicapped compared to the mobility of animals.  But what they lack in mobility, they more than make up for with their ability to make they own food from sunlight by photosynthesizing. And with the energy that sunlight provides, plants can create the food—such as pollen, nectar, and fruit—that entices insects and other animals to help them reproduce.  So how do plants protect themselves without fleeing from their predators?  That’s what Light Eaters is about.

I don’t know the source of this photo. It was sent to me in an email by someone who found it on Facebook.

How do plants perceive threats and react to them?

Plants can sense that they are being attacked by an insect in a variety of ways.  They can sense the vibration of the chewing, which is closely related to how animals hear.  The attack can also trigger an electrical impulse which can travel throughout the entire plant. 

Plants emit chemicals in response to the attack on their leaves and roots. The chemicals can repel the insect by making the plant unpalatable.  In a sense, the plant is producing its own pesticide, which has the potential to replace synthetic pesticides. 

The chemicals are also wafted into the air to serve as warning signals to their plant neighbors, who can then produce their own chemicals in preparation for attack. Some plants can distinguish between an attack that threatens individuals and those that threaten the entire community. They can tailor their warning messages accordingly, to send messages only to their relatives or to the entire plant community.  When plants are sprayed with herbicides, these chemical messages are masked by herbicides. (2)  Likewise, pollution can also muddle the chemical messages of plants and reduce their ability to perceive and respond to threats. (3)

Plants sometimes demonstrate a preference for their relatives in other functions as well.  They can make room for the roots of close by relatives and move branches to avoid shading their relatives.  They can also vary these accommodations depending on available resources, making room when there is plenty of water, nutrients, and light, but not when there’s not enough.

Such warning signals can also be sent via the underground root network, which connects plants in a community to one another through the network of mycorrhizal fungi that attach themselves to plant roots.  That network is also used by the community of plants to share resources, such as moisture and carbohydrates produced by photosynthesis.  The fungal network enables both communication and sharing of resources.  Herbicides that are carried to the roots of trees damage the fungal network, depriving trees of the nutrients they need to survive. (4) The widespread use of these herbicides by native plant “restorations” is one of many reasons why these projects rarely result in new landscapes of native plants. 

Can plants hear?

One of the first discoveries of the ability of plants to find what they need is the ability of tree roots to grow in the direction of water sources.  Mycorrhizal fungi attached to the roots of plants are clearly involved in guiding that connection.  Over 450 million years ago, the evolution of fungi enabled plants to move from water to land by delivering moisture from soil to roots of plants, greatly increasing abundance and diversity of plants. About 80% of plants today receive much of their nutrients and moisture through mycorrhizal fungi. (5)

Now there is evidence that plants may also be able to hear the sound of water to direct the growth of roots.  The researcher who made that discovery encased the roots of a plant in plastic pipe so that the roots could not sense the availability of moisture.  The plastic pipe formed a “Y” to give the roots the option of growing in one direction or the other.  The researcher played a recording of running water at the end of one pipe.  The roots grew in the direction of the recording of running water.  This is still a controversial discovery, because other researchers have found it difficult to replicate. 

The replication of breakthrough scientific discoveries is one of the ways that science moves forward.  It is a not a reliable method of confirming or rejecting a new discovery because there are always many variables operating simultaneously that are difficult to control, particularly in field studies, and researchers have rarely identified all the variables involved in the phenomenon they are observing.

The academic career of David Rhoades is an example of the dangers of being too far ahead of your academic colleagues and a reminder of the conservatism inherent in academic science.  Rhoades was a chemist at University of Washington and the author of a study that made the first report of warning signals that plants under attack send to their neighbors via volatile chemicals in the atmosphere. 

The forest on Rhoades’ campus was being killed by tent caterpillars.  He studied the spread of the caterpillars until the insect infestation was stopped by the chemicals that the unaffected trees infused into their leaves.  The chemicals killed the caterpillars and the spread of the insect in the forest was stopped.  Backed by a mountain of carefully accumulated data, Rhoades concluded:  “This suggests that the results may be due to airborne pheromonal substances!”

Rhoades was met with resistance to this new information from his colleagues.  Then he had trouble replicating his original study.  When his grant applications were rejected, he gave up.  He left academia and taught chemistry in a local community college to make a living.  Years later, other researchers figured out why he was unable to replicate his original study.  The airborne chemicals that trees produce are seasonal.   Rhoades’ original study was done in the spring and Rhoades was trying to replicate the study in the fall.  The scientists who eventually confirmed Rhoades’ finding did so in the laboratory where conditions are easier to control.

Plants collaborate with animals to protect themselves and reproduce

The Light Eaters reports many remarkable observations of interactions of plants and animals.  Here is a sampling of these stories:

  • If bumblebees emerge from hibernation before plants begin to bloom, the hungry bee bites the plant’s leaves to trigger the bloom that delivers the nectar the bees need.
  • Plants must use their limited resources to make pollen and nectar.  Some plants can ration the delivery of the pollen and nectar that attracts their pollinators by timing the delivery with the anticipated arrival of the pollinator.  The plant estimates the time of arrival of the insects based on its memory of past visits. 
  • Bats find the plants they pollinate by using echolocation sonar to locate them in the dark.  Some plants that are pollinated by bats have evolved saucer-like petals that act like a satellite dish to receive the sonar ping to help bats find them. 
  • Some corn, cotton, tomato and tobacco plants can emit chemical distress signals to summon tiny parasitic wasps to kill caterpillars such as tobacco budworm and corn ear worm.
  • Many orchids are pollinated by wasps.  Some orchids attract wasps by mimicking the chemical pheromones of the female wasp.  The orchid is pollinated by the attempt of the male wasp to mate with what he supposes is a female wasp.
  • Some plants form partnerships with ants by secreting a sugary substance that feeds the ants, who eat the insect predators of the plant. 

Can plants see?

The observation that plants are capable of mimicking animals and other plants is not new.  In the early 1900s, a Russian agronomist observed that weeds in food crops have sometimes mimicked the food crop and thereby evaded the hand-weeding that was the method used by farmers to eliminate competition for their crop.  Rye, oats, and lentils were initially considered weeds of wheat and rice.  Over time, they evolved the seed heads that qualified them as food crops. 

More recently, weeds that are killed by herbicides within crops that have been genetically modified to be resistant to the herbicide have engaged in mimicry at the biochemical level to also become resistant to the herbicide.  Those who engage in chemical warfare against plants do not seem to understand that it’s a war they can’t win because evolution will enable plants to develop resistance to their poison. 

Like many of the remarkable capabilities of plants, scientists can observe the phenomenon, but they are rarely able to explain the mechanism that makes it possible, beyond the evolutionary force of natural selection, which achieves a better adapted plant or animal through a series of mutations and genetic and epigenetic drift.  Each change in the species is a trial balloon.  If the change works, it’s a keeper.  If it doesn’t, it’s in the dustbin with some 99% of the estimated 5 billion species that have lived on Earth.  The dominant evolutionary force is random, irrepressible, complex change.  The notion that humans are capable of stopping evolution is absurd.

In 2014, a Peruvian ecologist discovered a vine in the Chilean rain forest that is capable of quickly taking on the shape of almost any plant that it grows beside.  Nicknamed the chameleon plant, many tests proved that the vine can mimic many different species of plants.  Presumably this mimicry enables the vine to become invisible in the sense that it blends in with whatever plants it grows amongst.  It’s a disguise, if you will, that protects the plant from its predators. 

The chameleon vine is able to mimic plants that are native to their locations as well as plants that are foreign to the region.  In other words, mimicry is not the result of a long evolutionary co-existence.  This finding is another blow to the nativist myth that plant and insect associations are the result of co-evolution that makes insects dependent on native plants.  The associations between plants and insects evolved long before the plants and insects moved into new regions.  Plants and insects retain that association as they change in response to their new environment and as the result of mutations and genetic drift. 

Until recently, there was a debate among scientists about how the chameleon plant morphs itself into an entirely different shape.  One school of thought speculates that plants have an organ that performs much like our eyes.  Another school of thought is that horizontal gene transfer (6) from the bacteria inhabiting the plant being copied to the plant doing the copying achieves this transformation. 

A study (7) published in 2022 seems to support the hypothesis that some plants have some type of organ that functions like our eyes.  The study found that the chameleon vine was capable of mimicking an artificial leaf.  The plastic leaf contains no chemicals or bacteria. 

In conclusion

The Light Eaters reports many other capabilities of plants that aren’t covered in this article.  If it’s a topic of interest to you, the book is well worth reading.  It’s well researched and well written.  It is also thoughtful because it asks us to ponder the philosophical question of whether or not this new(ish) knowledge of plants adds up to intelligence, consciousness, and agency.  Ms. Schlanger dodges that question by reminding us that there is not consensus agreement about what any of those descriptions actually mean.

Now we must add a few caveats that we hope will put this important topic into perspective:

  • Not every plant species has all of the capabilities described in The Light Eaters.
  • Those that do have such capabilities may not consistently use them because every plant is responding to a specific environment in a specific place.  Plants are inseparable from their environment.  A plant that has plenty of water and plenty of light behaves differently than plants with less resources.  Sweeping generalizations about plants are usually ridiculous.  For example, it makes no sense to claim that native berries are more nutritious than non-native berries. (8)
  • Plants have the potential to develop such capabilities, depending on their specific circumstances.
  • Without a brain or a nervous system, plants seem to organize a response to stimuli by functioning as a decentralized network.    

The Light Eaters says as much about science as it does about plants.  There are fads in science, just as there are fads in every human endeavor.  Presently, much scientific investigation of botanical phenomenon is focused on genetics, which has misled the public to underestimate the plasticity of plants and animals.  In fact, the genome of a species is a flexible repertoire, with many genes unexpressed until triggered by a change in the environment in which the plant lives.  For many characteristics of species, the environment is a more powerful influence than genes. 

Science is better at observing than it is at explaining.  Explaining requires speculation and academic science studiously avoids speculation.  The reader of scientific studies is often left in a quandary.  Conclusions are often a contradictory list of maybes with a plea for funding for further investigations. That’s one of many reasons why science journalism is important to the general public’s understanding of scientific issues.  Ms. Schlanger goes out on a limb for us by speaking in comprehensible terms that many scientists refuse to use.  Thank you, Ms. Schlanger, for helping the public understand the plant world.


Shortly before publishing this article and after I had drafted my article, I received the following review of The Light Eaters from Arthur M. Shapiro, Professor Emeritus of Ecology and Evolution, UC Davis.  He has given permission to add his review to my article.
– Conservation Sense and Nonsense

Elizabeth Kolbert has a collective review of Schlanger and two other, similar books–“The Nation of Plants” by Gregory Conti and “Planta Sapiens” by Calso and Lawrence–in the new NY Review of Books (Oct.3). Her review is only lightly snarky because it’s clear she doesn’t know quite what to make of the “plant neurobiology” fad.

“When I read Schlanger (I haven’t read the others) I dug back into my library to find my copy of “The Secret Life of Plants” by Peter Tompkins and Christopher Bird (1973). I doubt that Kolbert realizes that the current fad is a rerun of the 70s!  Unlike Schlanger and perhaps the others reviewed by Kolbert, Tompkins and Bird is packed with overt woo-woo and makes little attempt to be “science-based.” The frank woo-woo is very 70s. But the underlying motivation for both waves is the same: philosophical panpsychism, the notion that consciousness is ubiquitous in Nature.

“There is nothing in the actual data discussed by Schlanger that obliges one to embrace panpsychism. The main reason to do so is that one WANTS to. That is, for some (many?) people it is very reassuring to believe that at least the biosphere, if not the entire universe, is sentient. (This has resonances with the Gaia Hypothesis.) This notion is an integral part of a number of cultural cosmologies, of which the most familiar to most Americans is probably Native American, broadly speaking. In the 70s many hippies embraced the Native American notions of “tree people,” “stone people,” etc. Some still do.

“Remember that I have taught community ecology for some 50 years, with an emphasis on coevolution. Things like inducible anti-herbivore defenses (chemical or morphological) and communicable defensive messages (plant pheromones, if you will) come as no surprise. Rather, they are predictable consequences of natural selection: if something can evolve, it probably will.  There is no logical necessity to invoke intelligence or consciousness to account for them. If you want to, go right ahead. But don’t call it science!

“I have never had a chance to pull up a mandrake plant. In the Middle Ages it was widely believed that if you did it would shriek, and the sound if heard would drive one mad. Thus one must cover one’s ears when doing so. Now, that is framed as a testable hypothesis!

“Are you familiar with the walking fern? If not, Google it. I am very fond of it, but never for a moment would I claim it has the property of wanderlust.

Arthur M. Shapiro, Professor Emeritus of Ecology and Evolution, UC Davis


  1. Zoë Schlanger, The Light Eaters: How the Unseen World of Plant Intelligence Offers a New Understanding of Life on EarthHarper Collins, 2024. The Light Eaters is the source of information in this article unless otherwise noted.
  2. Behrend*, J.E., & A.L. Rypstra (2018) Contact with a glyphosate-based herbicide has long-term effects on activity and foraging of an agrobiont wolf spider.  Chemosphere 194:714-721   doi: 10.1016/j.chemosphere.2017.12.038
  3. “Polluted Flowers Smell Less Sweet to Pollinators,” New York Times, February 16, 2024
  4. K. Hage-Ahmed, “Arbuscular mycorrhizal fungi and their responses to pesticides,” Pest Management Science, September 25, 2018
  5. Thomas Halliday, Otherlands, A Journey Through Earth’s Extinct Worlds, Random House, 2023
  6.  Conservation Sense and Nonsense, “All Life on Earth is Related
  7. Jacob White and Felipe Yamashita, “Boquila trifoliolata Mimics leaves of an artificial plastic host plant,” Plant Signaling Behavior, 2022
  8. Conservation Sense and Nonsense, “Baseless Generalizations in Doug Tallamy’s Nature’s Best Hope”